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Climate Change Coverage in British
Kjersti Skaaraaen Herberg
Gjøvik videregående skole
This essay is an investigation of how conservative British tabloid and quality
newspapers skew the issue climate change. Through media analysis it discusses the research
question “to what extent do the British newspapers
The Daily Mail and
The Daily Telegraph
differ in the ways in which they deal with the issue of global climate change?
Twenty-nine articles from three weeks’ newspapers have been closely analyzed with
special respect to the four categories theme, purpose, point of view and target group, in order
to find a pattern in how climate change is presented. This has been organized to allow the
development of an argument, supported by key citations. An attempt has then been made to
gain insight in the reasons for the editorial decisions that the findings are a result of.
The main observation of the investigation is the sceptical line taken, as both
newspapers raise more doubt about the existence of climate change than up-to-date research
supports. They consistently take the average citizen’s point of view in matters concerning
expenses of climate change policies, and such expenses are the focal point of especially The
. The latter also focuses more on morally ambivalent celebrities and uses
exaggeration more extensively than The Daily Telegraph
. These observations are mainly
explained by news values, British culture, science skewing and target group.
The conclusion is that despite the main differences that The Daily Telegraph
sceptical than The Daily Mail
and that the latter oftener connects climate change to other
issues such as personal finances and celebrities, the two newspapers deal with global climate
change rather similarly. Although only two newspapers have been considered and only for a
short period, thus making a broad generalization inappropriate, the investigation gives an
impression of conservative British newspapers’ coverage of climate change.
Table of Contents
Part I: Analysis of the newspapers’ coverage of climate change
Media is often referred to as ”the fourth estate”. This demonstrates the strong position
it has taken in the modern society, where it both reflects and influences culture to a large
extent. Partly therefore, I have chosen to base my extended essay on newspaper investigation.
Another wish was to combine newspaper investigation with the issue global climate
change, which has become very important to our society. It is now a commonly accepted
facthat global warming will entail several serious consequences if we do not take action
Besides, climate change is a direct consequence of the materialistic culture that
characterizes the Western world today. When choosing a country for my research, Great
Britain stood out as an interesting choice for several reasons. Firstly, the country has taken
the lead internationally by being the first to set legally binding targets for greenhouse gas
emissions, and the awareness of climate change is very high, being 98%Six of ten Britons
are worried about climate change3, so I assumed that this concern would be reflected in the
newspapers. Secondly, it has historically been one of the main pollutersdue to large
emissions from early on. A third reason for my choice was the fact that British people are
among the most avid newspaper readers in the world. 82.0% of all British adults read a
newspaper every day, so newspapers are an important source of information.
In order for my project to comply with the word limit and time available, I have
chosen to limit my investigation to two newspapers: The Daily Telegraph
and The Daily Mail
Firstly, I chose these two because the first is a broadsheet and the second a tabloid newspaper,
1 See appendix 1.
2 IPCC Working Group II Report "Impacts, Adaptation and Vulnerability", http://www.ipcc.ch/ipccreports/ar4-wg2.htm, accessed 12.06.09, 09:57. Technical Summary.
3 Department for Environment, Food and Rural Affairshttp://www.defra.gov.uk/environment/climatechange/uk/individual/attitudes/, accessed 11.06.09, 13:364 Singer, P. One World: The Ethics of Globalization
. (New Haven and London: Yale University Press, 2004),Chapter 25 The Newspaper Society : http://www.newspapersoc.org.uk/Default.aspx?page=897, accessed 09.06.09, 13:23
and published research has shown a sharp distinction between the two categorieIt was my
opinion that when considering one difference, the other factors should be as equal as possible.
Therefore, I chose two newspapers that are both considered to be conservati
As a result of these choices, I ended up with the research question “To what extent do
the British newspapers The Daily Mail and The Daily Telegraph differ in the ways in
which they deal with the issue of global climate change?”
My intention is to study the
newspapers, mainly by applying my knowledge of analysis, but also by comparing it to
published research on media’s coverage of climate change and on the differences between
6 See appendix 2.
7 BBC News: http://news.bbc.co.uk/2/hi/uk_news/3409185.stm, published 19.01.04, accessed 10.06.09, 09:188 The Guardian: http://www.guardian.co.uk/media/2001/feb/22/dailymail.pressandpublishing, published22.02.01, accessed 10.06.09, 10:36
In order to answer my research question, I will analyze the news articles about climate
change printed in my two chosen newspapers over a time period of the three consecutive
weeks 17-19 in year 2009, including Sundays. The requirement I have set for the articles is
that the on-going global climate change is directly linked to the main theme. My focus is on
news articles, but editorials and commentaries, and also articles where climate change is
discussed to a smaller extent, may be included where it is relevant.
My task is then to analyze the texts with respect to topic, purpose, point of view and
target group. I do not find it suitable to compare numbers, because length of the articles as
well as the newspapers’ total number of articles may vary. Through a qualitative analysis, I
will be able to get an overall impression of how the issue climate change is dealt with. Then I
will try to give reasons for my findings by relating them to climate change research, the
newspapers’ values, political allegiance, readership, news values and British culture. I should
by this become able to reach a conclusion on how the newspapers skew the issue global
climate change, and what reasons they may have to do so.
The Daily Telegraph
is Britain’s best-selling quality daily newspaperwith a
circulation of 2,200,000 every daIt is part of the Telegraph Group, owned by
The Daily Mail
is the second-largest tabloid paper with a
circulation of 820,000 a day4, and is owned by Daily Mail and General Trust. Both
newspapers target the middle- and upper-middle class of educated adult
9Telegraph Media Group: http://advertising.telegraph.co.uk/static/KeyFacts.aspx, accessed 10.06.2009, 10:1910 Daily Mail and General Trust pcl:http://www.dmgt.co.uk/corporatestructure/anmediaassociatednewspapers/circulationfigures, accessed10.06.2009, 10:1711 J. Price, J. Nicholas: AS Media Studies
(Cheltenham, UK: Nelson Thornes Ltd, 2003), Chapter 7
Part I: Analysis of the newspapers’ coverage of climate change
The texts that were found relevant for analysis with respect to the research question,
counted 13 in The Daily Mail
, including 12 news articles and one commentary, and in The
the corresponding figures are 13 and three. Climate change was not once on
the front page. The results of the analysis are as follow:
The Daily Mail
: The most striking observation is The Daily Mail
’s large focus on what
negative short-term consequences Britain’s binding emission targets will have for the average
citizen, including personal finances and having to change one’s lifestyle. Out of 13 articles,
five were mainly about tax increases. We are given the impression that being able to combat
global warming requires too large an effort to be feasible.
For example, taxes are the focus of an article on April 23rd:“Tough new targets to
tackle climate change will cost every household at least £600 a year, push more than a million
into poverty and send fuel bills soaring. […] The targets will be legally binding, even if every
other country in the world continues to increase carbon emissions.” This appeals to people’s
sense of justice, because it seems unfair that some countries should do more than others that
On April 21st, Matthew Elliott, Chief Executive of the TaxPayers’ Alliance, is quoted
about the Government’s emission targets: “The Government is running a serious risk of
appearing to fiddle while Rome burns. Introducing green policies that will cost billions at a
time when the economy is in crisis is frankly irresponsible.” His job is obviously to be
negative towards tax increases, not to be concerned about the global future. We are
convinced that we need not hurry, and that right now we have not got the capacity to think
12Daily Mail Connected MyLife Survey: http://www.dailymailconnected.co.uk/pdf/news_mylife.pdf, accessed10.06.09, 11:37
about anything but ourselves. April 24th, the Mail
had an article about carbon capture and
storage at coal plants, telling us that this might just be wasted money: “[Ministers] admit the
technology has not been tested on a large scale, will cost billions of pounds to implement, and
could add around two percent to household electricity bills.” On May 5th, the Mail
similar article with the heading “Miliband’s global warming law ‘could cost £20,000 per
family’”. By close-reading the rest of the article, one understands that this sum is to be paid
over a period of 41 years. The journalist focuses on the costs to taxpayers, rather than the
benefits this will have for the environment.
April 21st, the Mail
printed a comprehensive and negative article about wind power:
“Wind power is a catastrophe, threatening to despoil our beautiful countryside as an
increasing number of hills and mountains succumb to regiments of giant white windmills.
[…]There are already plans to build huge wind farms across some of Britain’s most glorious
landscapes.” This is an example of egoism and short-termism, since a major climate change
will most likely change landscapes a lot more than building wind farms would, e.g. through
April 23rd: “Critics said the targets would cost the economy £14 billion a year by 2020
and would have only negligible impact on climate change.” The highly controversial climate
change sceptic Bjørn Lomberg is cited as an expert, although he represents a very small
minority of environmental scientists: “It will cost billions of pounds and the net effect will be
to reduce world temperatures by one three-thousandth of a degree by the end of the century.”
These articles’ common message is that we can just as well give up combating global
warming, because it is too challenging.
13 Chapter 10.4 in IPCC, 2007: Climate Change 2007: Impacts, Adaptation and Vulnerability. Contribution ofWorking Group II to the Fourth Assessment Report of the Intergovernmental Panel on Climate Change, M.L.
Parry, O.F. Canziani, J.P. Palutikof, P.J. van der Linden and C.E. Hanson, Eds., Cambridge University Press,Cambridge, UK, 7-22
The Daily Telegraph:
The negative sides of combating climate change were the main topic of
only one article and an important topic of a second in the Telegraph
April 30th: “Meeting the target to cut greenhouse emissions by 80 percent by 2050 will
cost around £17 billion a year, or add around £700 on the average electricity bill by 2050.”
These are the total costs, so on average it gives a cost of £17 per year – not that much to make
a fuss about. May 9th: “After all, talking about fighting climate change is a bit of a nonsense
if the only practical way to get around rural areas is in a car.” The message to the readers is
that it is not worth combating climate change if it is unpractical and requires a lowering of
The Daily Mail
: Celebrities are the second major focal point of the Mail
, and especially
environmental spokesmen’s profligate lifestyles – this is the topic of five articles. On April
26th, the Mail
printed two articles about Prince Charles’ flying a private jet on his tour to
promote environmental issues, and on May 4th there were two similar articles about the
Foreign Secretary and Prime Minister: “Critics accused the pair of seeking to ‘live a life in
luxury’ at taxpayers’ expense – and pointed out that using private charter flights dramatically
The Daily Telegraph
: The Telegraph
printed three articles about celebrities, only one
focusing on their ambivalent moral, and two about Prince Charles’ environmental efforts.
The broad consensus among scientistaken into consideration, both newspapers take a
sceptical line on climate change. In fact, on April 26th, The Sunday Telegraph
wrote: “In his
book, [Lord Stern] criticises the media for giving any space at all to [the deniers of climate
change], when ‘the balance of logic and evidence is 99 percent or more to one’”.
The journalists often seem to throw doubt upon official figures, politicians and
scientists. They also tend to exaggerate facts to make global warming seem beyond common
sense, and to ridicule experts and environmentalists.
The Daily Mail:
May 5th: “the Government is committed to cut carbon emissions, blamed for
global warming”. This wording implies that the connection between greenhouse gas
emissions and global warming is not known, but merely thought, to exist. “[Mr Miliband]
denied the figures were framed to produce a convenient answer.” “Ministers insist the costs of
not acting on climate change would be higher than the price of acting now”. Both quotes
show that the journalist serves us the official version, but intimates that she herself does not
believe it. This is done by emphasizing that she is referring to official announcements, not
presenting personal or the newspaper’s views.
May 8th: “If we have a wet summer, it’s the fault of ‘global warming’. If we have a
rare fine summer, Gordon Brown takes the credit.” This gives the impression that climate
change is all about natural coincidences that are used by politicians to further themselves.
The Daily Telegraph:
A larger portion of the texts express scepticism, for example by using
the wording “supposed climate change”, as on April 29th. The Sunday Telegraph,
In an article with the headline “Save the planet rhetoric reaches crazy heights”, general
climate change research is referred to as “environmentalist propaganda”. The journalist
quotes environmental spokesmen saying that “failing to ensure one’s home is ‘energy
efficient’ was a ‘moral crime’, as ‘socially unacceptable’ as drink driving”, and that
“opposing wind farms should be as ‘socially unacceptable’ as not wearing a seatbelt”. The
above quotes have been taken out of their original context, and since one often expresses
oneself differently written and orally and the humour with which something might have been
said is not as easily caught when read, the quotes are made to seem much exaggerated and
rather ridiculous. This spreads distrust towards experts and pictures climate change as far
beyond common sense. The finishing paragraph is “Fortunately, the latest available data
show the downward trend in global temperatures continuing. One thing we don’t need to
worry about, it seems, is global warming.” This is just the journalist’s statement, and no
effort is made to back it up with data. Journalists’ tendency to fail to do so is also observed in
an article on British environmental reporting: “in both the tabloids and broadsheets, there was
little evidence provided, in the form of data, to substantiate the claims being made.”
Although they fail to support their own claims with arguments, the journalists often try
to rebut experts’ claims by finding other data:
The Sunday Telegraph
April 26th: “the Stern Review went out of its way to cherry pick the
most alarming possible predictions about the impacts of climate change and then to
exaggerate them still further.” This argument is backed up by figures from other experts that
disagree with Stern, underlining the message that climate change is grossly exaggerated, if
even true. In the finishing line, Stern is even described as “lost in his apocalyptic dreams”.
The same is done in a similar article about Stern in The Sunday Telegraph
on April 21st. The
journalist backs up many of her arguments with figures that differed from Stern’s – only to be
forced to apologize due to misinterpretations a week later.
There are also other examples where quotes are taken out of their original context to
be exaggerated: April 21st there were two articles about Lord Stern’s warnings on climate
change. One told that “he said […] Florida and Bangladesh could disappear, alligators could
live at the North Pole and millions of people would have to migrate.” However, he is quoted
15 N. Taylor and S. Nathan: “How Science Contributes to Environmental Reporting in British Newspapers: ACase Study of the Reporting of Global Warming and Climate Change”, The Environmentalist
, December 2004,doi: 10.1023/A:1020762813548
directly in the second article: “The last time the world was 4-5 degrees above where we are
now was 30-50 million years ago, when much of the planet was swampy forest and there were
alligators near the Pole”. There is a rather large difference between the two, and one might
suspect that the journalist has misinterpreted Stern deliberately to make his words seem too
over-dramatized. In The Sunday Telegraph
April 26th, another journalist goes even further:
“Lord Stern predicts that global warming will make the Artic an ideal habitat for alligators”.
The article, where Stern is referred to as “the world’s Scaremonger-in-Chief”, furthers my
argument that data and quotes are manipulated.
Often, humour and self-irony are used to forecast a future with global warming. This
has the effect that it suppresses the gravity of the situation. It conveys the message that it is
better to stay calm, think about oneself today and not worry unnecessarily. May 2nd: “English
lawns will become a sign of ‘social and moral decadence’ as climate change takes effect in
the next decade, say horticulturalists. […] Gardeners will have to plant more Mediterranean
On May 10th, The Sunday Telegraph
had the headline “The elements conspire against
the warmists”. “The frenzied efforts of the warmists to panic us over all that vanishing Arctic
and Antarctic ice are degenerating into farce”. Later, the journalist refutes Ban Ki-moon’s
claim that the polar ice is melting quickly by using contradicting figures from the US National
Those who are concerned about global warming are often mocked and made fun of,
and are referred to in terms as “romantic greens” (the Mail, April 21st) and “crazed eco-
warriors” (the Telegraph, April 21st). This makes it less alluring to believe and sympathize
Although both newspapers are sceptical at times, most articles in the Mail
and some in
acknowledge the threat and the necessity of action. For instance, the Mail
refers to “the two great crises of our age – climate change and the energy gap” (April 21st) and
“harmful greenhouse gases” (April 19th). On May 10th, the Telegraph
“politicians should force us into setting a concrete example to the developing nations on
climate change” and of course both quote experts supporting the existence of climate change.
Very often, the Mail
exaggerates global warming, so that it seems a nearly impossible
task. An article in the Sunday Mail
on May 3rd states that “anyone who eats meat regularly
dramatically increases the size of their carbon footprint.” Although this might well be true,
the article treats it in such a way that we get the impression that everything we do is wrong,
and that combating climate change is far too large a task to master. Simon Retallack, head of
the Institute for Public Policy Research, says: “The public become disempowered because it’s
too big for them; and when it sounds like science-fiction, there is an element of the unreal
there.”16. I have not observed this tendency to the same extent in the Telegraph
None of the newspapers mention “small actions” or what individuals can do for the
climate. This is also observed by Solitaire Townsend, MD for the sustainable development
communications consultancy Futerra: “The style of climate change discourse is that we
maximise the problem and minimize the solutiAnother finding that is common for both
papers is that they only write about Britain – British climate policies, experts, celebrities, etc.
16 BBC news, 21.04.09. Accessed fromhttp://news.bbc.co.uk/2/hi/science/nature/5236482.stm, 15.06.09, 15:28
Both newspapers focus largely on the costs to the individual of actions to combat
climate change, but The Daily Mail
does so to a significantly larger extent than The
Many of the articles feature celebrities, but this occurs more often in the Mail
Both newspapers throw doubt upon the existence and gravity of climate change, but the
more than the Mail
. Scepticism is aroused by trying to rebut experts,
ridiculing environmentalists, exaggerating and using humour.
Most often, climate change is acknowledged a threat.
has a tendency to exaggerate.
The focus is held on Britain, although it is a global topic.
Part II: Evaluation of the findings
Having analyzed the texts, summed up findings and considered the effects of them, it
remains to gain an understanding of why
these decisions have been made, to fully answer the
Instead of considering what benefits humanity and the society as a whole, the
newspapers most often choose to see things from the individual’s point of view. Both
papers target aspiring middle-class adults, who are likely to be money-conscious and
alert to what the tax money is spent on. Consequently, the readers disapprove of
increases in personal expenses – especially when they are spent on things that will not
benefit them immediately, as is the case in global warming. This interests and arouses
them and thus makes it valuable news. Perhaps the target groups can explain why this
is the focus of a larger proportion of the articles in the Mail
, than in the Telegraph
Although the readerships of both are claimed to be rather wealthy adults, there is a
tendency that tabloids sell better among downmarket readerand these have more
reason to be concerned about personal finances. Another explanation can be the
differences in newspaper categories, since tabloids often focus more on the lives of
Celebrities selPeople like hearing about seemingly successful people’s mistakes
and faults, because knowing that no-one is perfect legalizes our own faults. Besides, it
is annoying to readers that politicians urge people to lower their level of luxury, when
17 J. Price: GSCE Media Studies
(Cheltenham, UK: Nelson Thornes, 2003), Chapter 818 See appendix 2.
19 P. Rayner, P. Wall, S. Kruger: Media Studied: The Essential Resource
, Part 3: Media Institutions. (London:Routledge, 2004)
not doing so themselves. Negative news sell better than positive newsbecause
people find it more interesting to read about what people do wrong, than what they do
right. The Royalty gets large publicity, because Great Britain is a monarchy where the
Queen is a symbol of national unity, identity and priand people are interested in
what the royal family does. It is not surprising that the Mail
more than the Telegraph
, because this is characteristic for tabloi
One reason the newspapers give space to sceptics, is their effort to balance the news.
The Boykoff brothers have done research on the US quality press' skewing of climate
change, and concluded that their “adherence to balance actually leads to biased
coverage of both anthropogenic contributions to global warming and resultant
actiThis is likely to be the case in Britain, too. Especially the Telegraph
itself on being an enquiring newspapeand therefore tries to rebut experts and find
The scepticism might to a certain extent be explained by the newspapers’ political
allegiance, since both are conservative. Although both the Labour Party and the
Conservative Party acknowledge the necessity of action in the energy policy papers,
Labour suggests significantly more comprehensive policies. Labour has set specific
goals for CO2-reduction and the provision of renewable energy, whereas the
Conservatives focus more on adaptation to climate change and the difficulties in
20 A. Bell, M, Joyce, D. Rivers: Advanced Level Media
, (London: Hodder Arnold, 2001) Chapter 7: News21 The Daily Telegraph
, http://www.telegraph.co.uk/news/newstopics/theroyalfamily/2122182/Britain-should-get-rid-of-the-monarchy-says-UN.html, published 13.06.08, accessed 11.06.09, 12:3422 See appendix 2.
23 M. and Jules Boykoff: “Balance as bias: global warming and the US prestige press”, 2003,doi:10.1016/j.gloenvcha.2003.10.00124 J. Price, J. Nicholas: AS Media Studies
(Cheltenham, UK: Nelson Thornes Ltd, 2003), Chapter 7
providing enough cheap energy, rather than combating global warmingThis
suggests (but does not prove) that the Conservatives are less worried about global
warming, although it does not necessarily apply to the readers in general. Still, readers
might be thankful for sceptical arguments, because this serves an excuse to be passive
and keep lifestyles unaltered. Seeing the scepticism in relation to the reluctance to
change living standards is also in accordance with an article on British newspapers’
reporting on climate change, which concluded that there was “made little attempt to
address the suspected causes that would inevitably involve criticism of highly
So, although it is a rather unexpected finding that a quality newspaper is more
sceptical than a popular newspaper according to the research that quality newspapers
are more concerned about facts and scientific correctneit may be explained by
their wish to take the readers’ side, and to sell themselves as enquiring.
As mentioned, most Britons are concerned about climate change. The broad
consensus among scientists, the high level of education and the free information flow
taken into consideration, it would practically be impossible for a large newspaper to
Media professor Roy Greenslade claims that the Mail
has a consistent tendency to
exaggerate: “The Mail
presents every facet of life in as gloomy and threatening a light
25 The Labour Party’s environment policy paper: http://www.labour.org.uk/environment, accessed 18.06.09,09:4326The Conservative Party’s energy policy paper,http://www.conservatives.com/Policy/Where_we_stand/Energy.aspx, accessed 11.06.09, 13:4527 N. Taylor and S. Nathan: “How Science Contributes to Environmental Reporting in British Newspapers: ACase Study of the Reporting of Global Warming and Climate Change”, The Environmentalist
, December 2004,doi: 10.1023/A:102076281354828 See appendix 2.
as possible. […] Playing to the fears and narrow-mindedness of its audience, it
magnifies their xenophobia and hypochondria, panders to their envyheir
tendency to exaggerate arouses readers, making them interested in reading the articles.
Proximity is one of the twelve news values identified by Galtung and Ruso this
explains why the newspapers keep their focus on Britain.
We see that many of the newspapers’ decisions can be explained by news values, science
skewing, target group and aspects of British culture.
29 Roy Greenslade quoted in J. Price, J. Nicholas: AS Media Studies
(Cheltenham, UK: Nelson Thornes Ltd,2003)30 Reference to Johan Galtung and Mari Ruge in A. Bell, M, Joyce, D. Rivers: Advanced Level Media
The conclusion is that although there are some significant differences, The Daily Mail
and The Daily Telegraph
deal with the issue global climate change rather similarly, and that
’s coverage seems more sceptical towards the existence of climate
change than the Mail
’s, although it is the case for both that the proportion of sceptical articles
is larger than the proportion of scientists that are so. This must be seen in connection both
with target group and political allegiance. The Telegraph
has an enquiring approach,
supposedly because that is the way it wishes to sell itself. The Mail
’s tendency to exaggerate
may also partly be explained with the newspaper’s general journalism.
Both newspapers take the individual British citizen’s point of view in matters like tax
increases, because this is coherent with the reader’s opinions and interests. However, this is
to a greater extent the case in the Mail
than in the Telegraph
, possibly due to different target
groups and newspaper categories. Another significant observation is that especially the Mail
focuses largely on celebrities, presumably due to news values, the newspaper category and, to
These findings have been shown likely to be the result of conscious journalism and
intentional editorial decisions. Since the essay only considers two newspapers over a limited
time-period, generalizing the findings to apply to all conservative broadsheet and tabloid
newspapers is inappropriate. Nevertheless, the investigation serves as an indicator of British
newspapers’ coverage of climate change.
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The distinction between tabloid and broadsheet (quality) newspapers refers to formats
in which they are printed. However, there are often other large differences as well, both in
form and content. There are of course large variations, and the stereotypes do not fit for all.
Still, I will list some characteristics that can often be observed:
Source: Bell, A., M. Joyce, D. Rivers. Advanced Level Media
. London: Hodder Arnold, 2001. Chapter 7: News
Michael Anderson is a registered Psychologist in private practice in Torquay, Victoria. He has been practicing mindfulness meditation for over 25 years. In 1990 he became increasingly interested in Buddhism and mindfulness practice, and has from this time worked to integrate Western psychology and mindfulness approaches. He has training in ACT, Without a personal practice
Chem. Res. Toxicol. 2006, 19, 164-172 The Greater Reactivity of Estradiol-3,4-quinone vs Estradiol-2,3-quinone with DNA in the Formation of Depurinating Adducts: Implications for Tumor-Initiating Activity Muhammad Zahid, Ekta Kohli, Muhammad Saeed, Eleanor Rogan, and Ercole Cavalieri* Eppley Institute for Research in Cancer and Allied Diseases, Uni V ersity of Nebraska Medical Cen